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Fabricius nomad bee, Nomada fabriciana, a small red-and-black cuckoo bee on bare ground
Nomada fabriciana, the Fabricius nomad bee. Janet Graham, CC BY 2.0, via Wikimedia Commons.
Nomada fabriciana| Nomad bee (cuckoo) Common Red-abdomen

UK Bee Species

Fabricius Nomad Bee (Nomada fabriciana)

Nomada fabriciana (Linnaeus, 1767) · subfamily Nomadinae · family Apidae


Here is a small oddity of scientific naming: this bee was named to honour Johan Christian Fabricius, but not by him. Linnaeus, Fabricius's own teacher, described the species in 1767 and chose the epithet fabriciana as a tribute to his brilliant Danish pupil, then in his twenties and already at work on the vast insect catalogues that would fill his life.[3] Two and a half centuries later, a teacher's small gesture is still spoken aloud every time the bee is recorded.

The bee itself is a rust-orange, wasp-shaped cuckoo, only seven to ten millimetres long, hairless and long-legged: one of the earliest nomads on the wing each spring and one of the commonest in Britain. It builds no nest and gathers no pollen. Instead it slips into the burrow of a mining bee, lays an egg on the cell wall, and leaves its own larva to kill the host young and eat the pollen store left behind. See where it sits among Britain's bees on the UK Native Bee Species Map, or among the world's bees in the World Bee Atlas.

The bee at a glance

Scientific nameNomada fabriciana
AuthorityLinnaeus, 1767
FamilyApidae (Nomadinae)
SizeSmall, 7 to 10 mm
ColourRusty-red abdomen, black
LifestyleCleptoparasite (cuckoo)
ActiveMarch to August, two broods
Main hostAndrena bicolor
UK statusCommon, England & Wales
Full taxonomy
KingdomAnimalia
PhylumArthropoda
ClassInsecta
OrderHymenoptera
FamilyApidae
SubfamilyNomadinae
GenusNomada
SpeciesNomada fabriciana

A bee that lives like a wasp

Nomad bees are cuckoo bees, cleptoparasites of the mining bees in the genus Andrena.[4] A female Nomada has none of the pollen-carrying hairs of most other bees; she is hairless, wasp-shaped and shining, with a single job. She finds the burrow of her host species, slips inside while the owner is away, and lays her own egg on the wall of an unsealed cell.[4] The genus name is Greek for "wanderer", after that habit of drifting low over the host's nesting ground rather than digging a home of her own.[7] What happens next is where the wasp-like life gets its edge.

How a nomad takes over a nest

The four steps of cleptoparasitism, from break-in to inheritance

1 Slips in Enters the host's open burrow 2 Hides an egg Lays it inside the cell wall 3 Removes the rival The hatchling kills the host egg or larva 4 Inherits the larder Eats the pollen store the host gathered No pollen gathered, no nest built

One host above all others

Every nomad is tied closely to its host, and this one is no exception. Its main host across Britain and Europe is Gwynne's mining bee, Andrena bicolor, and where that small brown bee is common the Fabricius nomad is common too.[5] A handful of other mining bees are used more occasionally: hawthorn (A. chrysosceles), yellow-legged (A. flavipes), buffish (A. nigroaenea) and short-winged (A. angustior).[5]

One of the odder consequences is the huge size range of adults in a single population. It is thought to track the size of the pollen cell the larva grew up inside: a cell stocked for a small host produces a small nomad, one stocked for a large host a large one.[5]

Gwynne's mining bee, Andrena bicolor, the main host of the Fabricius nomad bee
Gwynne's mining bee (Andrena bicolor), the host this nomad depends on almost entirely.

Two waves a year

In Britain the Fabricius nomad bee is bivoltine, running two generations a year to track its equally bivoltine host.[6] Adults visit dandelions, buttercups and umbellifers for their own nectar, but never carry pollen home. Look for them not on flowers so much as on the ground, patrolling patches of bare, warm soil where the host's burrows are hidden.[1]

Spring brood
March to June. The first flight, chasing the spring generation of Gwynne's mining bee.
Summer brood
June to August. The second flight, tracking the host's summer generation.
Where the small brown Gwynne's mining bee nests, the red-and-black Fabricius nomad is rarely far away.

Where and when you will see it

The Fabricius nomad bee is widespread through England and Wales, especially in the south, and thins out further north with only sporadic records from Scotland and Ireland.[1] It turns up almost anywhere its host does, so nesting aggregations of Gwynne's mining bee on garden lawns, brownfield land, disturbed soil, chalk downland and open woodland are all good places to look.[1] It is also present across much of Europe from Ireland to Turkey, and across the Palaearctic further east.[6]

Telling it apart from other nomads

Britain has more than thirty species of Nomada, and many look alike; a firm identification is often a hand-lens job.[2] The Fabricius nomad has three features that, taken together, no other British nomad shows:

  • Banded antennae. Reddish, with a distinct black band on the middle segments, giving a striped look in the field.
  • Two-toothed mandibles. Bidentate jaws, visible under a lens.
  • Black labrum. The small plate above the mouth is black, not pale.

The abdomen is mainly reddish, sometimes with narrow paler bands at the segment edges. A larger, chunkier nomad with all-yellow legs is more likely to be Gooden's nomad bee or Marsham's nomad bee.[2]

Why a cuckoo bee is good news

Cuckoo bees can look, at first, like a drag on pollination: they gather no pollen, build no nests, and their presence means fewer host cells survive. In practice the story runs the other way. A cuckoo is only ever as abundant as its host, so a healthy nomad population is a signal that the mining-bee community beneath it is thriving.[7] A lawn that supports a red-and-black nomad also supports the small brown mining bee it depends on, and the wider web of ground-nesting bees around them, which is why ecologists have proposed cuckoo bees as useful indicator species for the health of a whole bee community.[8] See how the pieces of that web fit together in The Plate Without Bees.

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Frequently asked questions

What is the Fabricius nomad bee?
It is a small cuckoo bee, Nomada fabriciana, in the family Apidae. Seven to ten millimetres long, hairless and wasp-shaped with a rusty-red abdomen and reddish black-banded antennae, it lays its eggs in the nests of Gwynne's mining bee and a handful of related mining bees.
Why is it called the Fabricius nomad bee?
It was named by Linnaeus in 1767 in honour of his Danish student Johan Christian Fabricius, who went on to become one of the most productive entomologists of the eighteenth century. The word "nomad" comes from the Greek for "wanderer", after the female's habit of drifting over the host's nesting ground.
Is it really a bee, or is it a wasp?
It is a true bee, but it looks and behaves in some ways like a small wasp because it does not need pollen-collecting hairs and has evolved a hairless, brightly-marked body. All bees, including nomads, feed their larvae on pollen and nectar; the Fabricius nomad simply lets another bee gather the pollen for it.
What is a cuckoo bee?
A cuckoo bee is a species whose females do not build their own nests. Instead they enter the nest of another bee, called the host, and lay their eggs there. Their larvae eat the pollen store the host mother had gathered, usually after killing the host's own egg or young larva.
When are Fabricius nomad bees active?
They are bivoltine, with two generations a year. The first flight runs from March to June, chasing the spring brood of the host Andrena bicolor. The second runs from June to August with the summer generation. Almost any warm, sunny day in that window is a good time to look.
What is the main host of the Fabricius nomad bee?
The main host is Gwynne's mining bee (Andrena bicolor). Where Gwynne's mining bee nests in numbers, the Fabricius nomad is usually not far behind. Several other mining bees are used less often.
Do Fabricius nomad bees sting?
Very rarely. Solitary bees carry a sting but almost never use it. The Fabricius nomad has no colony or store of provisions to defend, is not territorial, and is completely safe around people.
Does the Fabricius nomad bee make honey?
No. Solitary bees, including cuckoos, do not make honey in harvestable quantities. Only the honeybee does. Compare the bee families in the World Bee Atlas.

Related species

Sources & references

  1. BWARS (Bees, Wasps & Ants Recording Society). Species account: Nomada fabriciana (Linnaeus, 1767): identification by reddish banded antennae, bidentate mandibles and black labrum, main host Andrena bicolor, wide English and Welsh distribution. bwars.com.
  2. Falk, S. Field Guide to the Bees of Great Britain and Ireland, Bloomsbury, and the Steven Falk Flickr collection: separation from other British Nomada by antennal banding and mandible shape.
  3. Linnaeus, C. (1767), Systema Naturae, 12th edition, original description. Etymology: the specific epithet fabriciana honours Johan Christian Fabricius (1745 to 1808), Danish entomologist and Linnaeus's pupil; Nomada from Greek for "wanderer" (NBN Atlas; GBIF).
  4. Rozen Jr, J.G. (1991). Evolution of cleptoparasitism in bees, with implications for their biology. American Museum Novitates (nomad-bee larval mandibles, egg placement in host cells, killing of host young). Also Odanaka et al. (2022) for phylogenetic host relations of Nomadinae.
  5. Falk, S. & Lewington, R. (2019). Field Guide to the Bees of Great Britain and Ireland (revised edition): main host Andrena bicolor, additional hosts A. chrysosceles, A. flavipes, A. angustior and A. nigroaenea, size variation with host.
  6. Crowley, L.M. et al. (2023). The genome sequence of Fabricius' Nomad Bee, Nomada fabriciana (Linnaeus, 1767). Wellcome Open Research 8:504 (Palaearctic distribution, UK status, bivoltinism, family placement).
  7. Else, G.R. & Edwards, M. (2018). Handbook of the Bees of the British Isles. Ray Society (Nomadinae biology, cleptoparasitism and the relationship between host abundance and cuckoo abundance).
  8. Sheffield, C.S., Pindar, A., Packer, L. & Kevan, P.G. (2013). The potential of cleptoparasitic bees as indicator taxa for assessing bee communities. Apidologie 44: 501 to 510.
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